Environmental Life Sciences (ELS) is a novel graduate degree program that will provide Ph.D.-level training in several complementary fields focused on interactions between organisms (plant, animal or microbe) and their environment. ELS will provide trans-disciplinary training that includes aspects of biology, geosciences, chemistry/biochemistry, environmental engineering, sustainability, social sciences, and mathematics. Graduate students will be trained in core classes, seminars, reading groups and research clusters. A focus will be collaborative and integrative study of the effects of environmental variation on fluxes of materials and energy across scales ranging from the organism to the globe. Our overall goal is to provide a unique Ph.D. degree program that produces students with a broad appreciation of environment-organism questions in the context of natural and anthropogenic environmental change.